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East Lawn Cemetery in Ithaca, New York, contains an aggregation of approximately 5.5 million Andrena regularis bees concentrated within a 1.5-acre area.
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Andrena regularis, commonly called the 'regular mining bee,' is a solitary wild bee species that nests underground and pollinates crops and wild plants.
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Rachel Fordyce, a lab technician at Cornell University, observed an unusual number of bees in East Lawn Cemetery during a walk in spring 2022 and collected specimens to show to Bryan Danforth.
Rachel Fordyce, lab technician
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"These are all over the cemetery," Rachel Fordyce told Bryan Danforth.
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Researchers estimate the bee population at East Lawn Cemetery ranges from 3 million to 8 million, with an average of 5.5 million individuals.
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The estimated bee population at East Lawn Cemetery is comparable to more than 200 honeybee hives.
Steve Hoge, undergraduate researcher
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"I'm sure there are other large bee aggregations that exist around the world that we just haven't identified, but in terms of what is in the literature, this is one of the largest," said Steve Hoge, lead author of the study.
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The study documenting the bee aggregation was published on April 13, 2026, in the journal Apidologie.
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Historical records show that Andrena regularis has been present at East Lawn Cemetery since at least the early 1900s.
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East Lawn Cemetery was established in 1878.
Bryan Danforth, professor of entomology
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"The research elevates the value of solitary ground-nesting bees and shows just how abundant these bees are, how important they are as crop pollinators, and that we need to be aware of these nest sites and preserve them," Bryan Danforth said.
Keven Morse, superintendent of East Lawn Cemetery
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Keven Morse, superintendent of East Lawn Cemetery, said he has seen deer, geese, hawks, foxes, coyotes, and countless bees during his family's 46 years managing the cemetery.
Keven Morse, superintendent of East Lawn Cemetery
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"And of course, bees, which he said have never stung him."
Keven Morse, superintendent of East Lawn Cemetery
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"I just felt bad having to mow in certain areas. There's probably three or four sections where they really migrate heavy, there's a lot of them," Morse said.
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Cemeteries provide good bee habitat because the land is peaceful, rarely disturbed, and largely free of pesticides.
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Approximately 75% of bee species are solitary ground nesters like Andrena regularis.
Bryan Danforth, professor of entomology
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"It's the most common lifestyle for bees," Danforth said.
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Female Andrena regularis create underground nests and place eggs in chambers filled with pollen and nectar.
Steve Hoge, undergraduate researcher
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"This species overwinters as adults, which is relatively rare, and that's part of the reason why they come up out of the ground so early in the spring, timed to the apple bloom," Hoge explained.
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In New York, Andrena regularis typically emerge in April when daytime temperatures regularly reach around 70 degrees Fahrenheit.
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Cornell Orchards, located about one-third of a mile from East Lawn Cemetery, may help support the bee population by providing abundant spring flowers.
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Andrena regularis prefers sandy soil, which East Lawn Cemetery contains in large amounts.
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Researchers used emergence traps—small mesh tents covering less than one square meter of ground—to estimate the bee population and study emergence patterns.
Bryan Danforth, professor of entomology
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"You capture a whole community of animals coming out of the ground with this approach," Danforth said.
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Between March 30 and May 16, 2023, researchers placed 10 emergence traps throughout East Lawn Cemetery.
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The traps collected 3,251 insects representing 16 species, with Andrena regularis overwhelmingly dominant.
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Male Andrena regularis emerge from the ground before females during warm periods in April.
Steve Hoge, undergraduate researcher
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"The males come out first and wait for the females, so that they have the best opportunities to mate and pass on their genes," Hoge said.
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The study documented brood parasitism by Nomada imbricata, a cuckoo bee species that lays eggs in Andrena regularis nests, where its larvae kill host larvae and consume stored pollen and nectar.
Bryan Danforth, professor of entomology
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"These populations are huge, and they need protection. If we don't preserve nest sites, and someone paves over them, we could lose in an instant 5.5 million bees that are important pollinators," Danforth said.
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Danforth and colleagues launched a global citizen science initiative to locate and protect ground-nesting bee aggregations.
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The study's co-authors included postdoctoral researchers Jordan Kueneman and Katherine Odanaka, undergraduate students Steve Hoge '24 and Cassidy Dobler '26, and lab technician Rachel Fordyce.
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Funding for the research came from the Cornell Atkinson Center for Sustainability, the National Science Foundation, and the Federal Capacity Funds program.
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